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Conifer seeds develop inside a protective cone called astrobilus.
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Sporangia may have formed a two-rowed spike orstrobilus.
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Fertile branches are produced in a terminalstrobilus(8.5 cm long), which is subtended by whorls of normally developed leaves.
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However, the two forms differ instrobilusmorphology.
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As with other cycads, it is dioecious, with the males bearing pollen cones (strobilus) and the females bearing groups of megasporophylls.
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In all cases, the lateral organs of thestrobilusare microphylls, bearing sporangia.
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